You might need to replace that fuse holder if the fuses test OK. I've seen them get a thin layer of oxidation that is impossible to see but interferes just enough with the fuses contact to make the game act up. The original factory fuse holder is hard to clean in my opinion and seems to be made of a metal that is not compatible with modern fuses and causes galvanization of the holder.
I guess if you really wanted to keep the original fuse holder, you could solder a new fuse holder atop the old so you can revert back to factory later.