I have had issues like this where the solution was either (1) the switch wire needed to be adjusted or (2) the cherry microswitch was just old or sticky and needed to be cleaned or replaced. In my cases it was the same deal - worked with in test mode but not gameplay. When the ball rolls over it’s different than when you press with finger. You can try putting a ball on the end of a magnet wand and pull it over switches manually to get a more realistic test, but even then the speed isn’t the same. It does seem odd that it is so many switches, though. In my case it was just a couple.

That is odd - I don’t understand how that could fix it. Root cause could be something else and somehow by messing around the cpu board it resolved. But glad your machine is working now.