Try re-pinning the j4 connector on the MPU board.
If u don't have the pins/tools to do it, try cleaning the header and extract the pins from the connector and clean them up with some 400-600 grit.
Start simple, then move on to more complex trouble shooting. Transistors, the U11 pia chip on the MPU board, bad sockets, fun stuff like that.
Most times, a good cleaning of the connector pins will do the trick temporarily until you can re-pin them.