Am I reading correctly that you have removed everything, including the LED OCD? If not, I would definitely suggest that first.
Disconnecting all the lamps and using the "testing with a jumper and bulb" section of that pinballrehab page is a good way to narrow it down between a board or wiring problem. Even better than a jumper is this test board:
http://www.siegecraft.us/presta/index.php?id_product=11&controller=product&id_lang=1
Often when installing LEDs, you can bump a socket and create a short. Occasionally, you can also get an LED bulb that has a short internally. That has bitten me twice now.