Those will work without issue.

As long as the circuit is running less than 32v.

If it's a 110v circuit, then no, a 32v fuse wouldn't be the proper rating.

Don’t confuse load with current.

The 20 amp fuse is for GI, which is at 7.3VAC.

I don’t know why they spec’d a 32 volt fuse for the 43 volt supply.

I think over time, fuses have become much more accurate about blowing at a certain amperage, not voltage. Blowing at or above a certain voltage, in my opinion, would require some type of voltage sensor or sparkgap. Way too expensive for a pinball machine.

The amperage is the critical part, I don't know why the voltage rating even needs to be specified, the fuse is a piece of glass with two copper ends with the fuse link soldered on each end, nothing is going to "arc over" if you put 110 volts on a 32 volt rated fuse.

Voltage rating on fuses is entirely independent of the value at which the fuse opens; it's still "this fuse has been manufactured to open at the rated current" regardless of voltage. The voltage rating is the maximum voltage that the fuse is designed to be used at, and has no real bearing on it's functionality. A 3 amp fuse rated at 250V is going to open at 3 amps, and a 3 amp fuse rated at 32V is going to open at 3 amps. The 250V fuse is just manufactured to be more robust, able to withstand being used in a higher voltage circuit, regardless of the current.