Keep in mind that there should be little that needs adjusting for switch settings unless someone before you has misadjusted switches.
The switches on L, M, N, Z1 and Z2 and all of the score reels need to be set up properly and actually have continuity when they need to. Yes, sixteen score reels worth. 3 of the 4 score reels have three functions: 1- at zero, 2- not at zero, 3- at 9 for carry over. The 10,000 reels do not have the carry over switches. The not at zero switches should be made when not at zero. This is the switch that drives power to the switch directly during reset, not through L, M, or N.
Setup of Z1 and Z2 are critical to get the reset sequence right.
Make sure each switch has good/positive wipe between the contact faces when it is the made position.
Sounds to me there is an issue in one or more of these.
You can exercise each relay manually to check function of each during start up and after start up.
If you really want to keep from distracting yourself pull the playfield totally out of the game. You should be able to get reels moved by actuating the L, M, and N relays for each player once the game has started up and score reels are all at zero.