Looking for help for my TOTAN. What might cause flippers to test good, but during game play, flipper buttons do not activate the flipper. I did a pcb swap out prior to this to problem solve another pcb issue, but this is new. I have double checked connections on mpu J212 and tried two different ribbon cables. Any advice would be appreciated!

Thanks for all your suggestions they did help me go back and recheck the already checked seatings. J 139 was off one pin, and after checking continuity on flipper optos, I put those back incorrect as well and fixed those too. Its a bitch being blind lmao. If you have good eyesight, thank the heavens above.