Video shows very strange operation.
Things I notice. Solenoids fire out of order in test mode. They are controlled by U11 but is often a connector issue.
Player 3 display is enabled at the wrong time (always?). This is controlled by U11 PA2 which also does lamp address 2 and switch strobe 2. PA2 shorted to something on PCB? Switch shorted to ground? Other displays are off and P1 does not rack up score staying at 00. Bad U10? Probably not the 4502 since there is problems beyond the displays.
U8 has an impossible date code. Does not mean it is bad, but could be suspect. Sounds like you swapped it out already tho.
Can also unplug the lamp board and isolate one display at at a time that has been verified working in another game in case problem on the lamp board or a display is fudging up the PIA through MPU j1. Check the zero X for 120hz and the display interrupt for 320-420hz as I suppose an interrupt problem is possible.
Seems likely the problem is at or around U10. The solenoids firing out of order probably separate connector issue between MPU j4 and driver j4.