A blown G.I fuse will test good in circuit because the meter will get a "good reading" the other way around through the transformer and lamps which are very low resistance. So you must remove the fuse to test it effectively.
Having said that, the entire G.I in this game is fused by one fuse. So if the playfield G.I is working the fuse is not the problem.
As MrBally stated, there are two sets of G.I circuit wires to the backbox that are connected together at each end. It would be a little unusual (though not impossible) for both circuits to fail at the same time. Pictures of the J3 connector on the rectifier board and also the block connector next to the backbox insert door will help.
You should check with your meter set on AC voltage if you measure around 7.8 volts AC at one of the actual lamp sockets that isn't working which will tell you if there's connector issues. On the chance you do get good voltage at a lamp socket it could be that all the backbox lamps are blown.